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Stanley H. Weitzman, John P. Wourms, South American Cyprinodont Fishes Allied to Cynolebias with the Description of a New Species of Austrofundulus from Venezuela, Copeia, Vol. 1967, No. 1 (Mar. 20, 1967), pp. 89-100
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Weitzman et al. (1967) studied this question.
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