Michel Gauthier-Clerc, Yannick Clerquin, Yves Handrich, Hyperinfestation by Ticks Ixodes uriae: A Possible Cause of Death in Adult King Penguins, a Long-Lived Seabird, Colonial Waterbirds, Vol. 21, No. 2 (1998), pp. 229-233
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Gauthier‐Clerc et al. (1998) studied this question.
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