Authors
Jon P. Costanzo, Michael F. Wright, Richard E. Lee, Jr., Freeze Tolerance as an Overwintering Adaptation in Cope's Grey Treefrog (Hyla chrysoscelis), Copeia, Vol. 1992, No. 2 (May 1, 1992), pp. 565-569
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Costanzo et al. (1992) studied this question.