Six species of Fragaria together with their subspecific entities are described and typified, and their chromosome number is given. A total of 392 collections have been investigated cytologically. So far as it seemed necessary for interpretation, historical notes are added. The following taxa are discussed: F. vesca (four subspecies, five forms, three cultivars), F. viridis, F. moschata, F. virginiana (four subspecies), F. chiloensis (four subspecies, two forms), F. × ananassa (two nothomorphs). New status is given to numerous taxa and two new taxa are described.
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G. Staudt (1962) studied this question.