Daniel J. Crawford, Tod F. Stuessy, Mario Silva O., Allozyme Divergence and the Evolution of Dendroseris (Compositae: Lactuceae) on the Juan Fernandez Islands, Systematic Botany, Vol. 12, No. 3 (Jul. - Sep., 1987), pp. 435-443
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Crawford et al. (1987) studied this question.
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