Ron Scogin, David A. Young, C. E. Jones, Jr., Anthochlor Pigments and Pollination Biology. II. The Ultraviolet Floral Pattern of Coreopsis gigantea (Asteraceae), Bulletin of the Torrey Botanical Club, Vol. 104, No. 2 (Apr. - Jun., 1977), pp. 155-159
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Scogin et al. (1977) studied this question.
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