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The difference in the levels of oleate and linoleate in the triglycerides of the related R84 and Illinois High Oil (IHO) maize (Zea mays L.) strains is determined solely by the In locus. A single locus controls the 20% difference in oleate and linoleate between C103 and R84. At least two loci control the levels of oleate and linoleate in C103 and IHO. Strong maternal effects are exhibited in the reciprocal F1's of C103 and R84. In contrast, no maternal effects were detectable in IHO ✕ R84 and C103 ✕ IHO.
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Roche et al. (1971) studied this question.