R. A. Sanderson, S. P. Rushton, A. J. Cherrill, J. P. Byrne, Soil, Vegetation and Space: An Analysis of Their Effects on the Invertebrate Communities of a Moorland in North-East England, Journal of Applied Ecology, Vol. 32, No. 3 (Aug., 1995), pp. 506-518
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Sanderson et al. (1995) studied this question.
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