Brian Moss, Vertical Heterogeneity in the Water Column of Abbot's Pond: II. The Influence of Physical and Chemical Conditions on the Spatial and Temporal Distribution of the Phytoplankton and of a Community of Epipelic Algae, Journal of Ecology, Vol. 57, No. 2 (Jul., 1969), pp. 397-414
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Brian Moss (1969) studied this question.