Eric T. Liknes, David L. Swanson, Seasonal Variation in Cold Tolerance, Basal Metabolic Rate, and Maximal Capacity for Thermogenesis in White-Breasted Nuthatches Sitta carolinensis and Downy Woodpeckers Picoides pubescens, Two Unrelated Arboreal Temperate Residents, Journal of Avian Biology, Vol. 27, No. 4 (Dec., 1996), pp. 279-288
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Liknes et al. (1996) studied this question.
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