F. J. Ebling, M. A. Sleigh, J. F. Sloane, J. A. Kitching, The Ecology of Lough Ine: VII. Distribution of Some Common Plants and Animals of the Littoral and Shallow Sublittoral Regions, Journal of Ecology, Vol. 48, No. 1 (Feb., 1960), pp. 29-53
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Ebling et al. (1960) studied this question.