Holger Deising, Ralph L. Nicholson, Marc Haug, Richard J. Howard, Kurt Mendgen, Adhesion Pad Formation and the Involvement of Cutinase and Esterases in the Attachment of Uredospores to the Host Cuticle, The Plant Cell, Vol. 4, No. 9 (Sep., 1992), pp. 1101-1111
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Deising et al. (1992) studied this question.