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Erich Scholtyseck, Datus M. Hammond, John V. Ernst, Fine Structure of the Macrogametes of Eimeria perforans, E. stiedae, E. bovis, and E. auburnensis, The Journal of Parasitology, Vol. 52, No. 5 (Oct., 1966), pp. 975-987
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Scholtyseck et al. (1966) studied this question.
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