A. D. Friend, F. I. Woodward, V. R. Switsur, Field Measurements of Photosynthesis, Stomatal Conductance, Leaf Nitrogen and δ13C Along Altitudinal Gradients in Scotland, Functional Ecology, Vol. 3, No. 1 (1989), pp. 117-122
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Friend et al. (1989) studied this question.
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