David B. Clark, Concepcion Guayasamin, Olga Pazmino, Cecilia Donoso, Yolanda Paez de Villacis, The Tramp Ant Wasmannia auropunctata: Autecology and Effects on Ant Diversity and Distribution on Santa Cruz Island, Galapagos, Biotropica, Vol. 14, No. 3 (Sep., 1982), pp. 196-207
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Clark et al. (1982) studied this question.
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