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N. S. Jones, D. J. Slinn, The Fauna and Biomass of a Muddy Sand Deposit off Port Erin, Isle of Man: With an Appendix on Methods Used for the Analysis of Deposits, Journal of Animal Ecology, Vol. 25, No. 2 (Nov., 1956), pp. 217-252
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Jones et al. (1956) studied this question.
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