J. W. Jones, H. B. N. Hynes, The Age and Growth of Gasterosteus aculeatus, Pygosteus pungitius and Spinachia vulgaris, as Shown by their Otoliths, Journal of Animal Ecology, Vol. 19, No. 1 (May, 1950), pp. 59-73
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Jones et al. (1950) studied this question.
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