J. Nathaniel Holland, Theodore H. Fleming, Mutualistic Interactions between Upiga virescens (Pyralidae), a Pollinating Seed-Consumer, and Lophocereus Schottii (Cactaceae), Ecology, Vol. 80, No. 6 (Sep., 1999), pp. 2074-2084
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Holland et al. (1999) studied this question.
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