James K. Roberts, Noelle A. DeSimone, Wilma L. Lingle, Leon Dure, III, Cellular Concentrations and Uniformity of Cell-Type Accumulation of Two Lea Proteins in Cotton Embryos, The Plant Cell, Vol. 5, No. 7 (Jul., 1993), pp. 769-780
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Roberts et al. (1993) studied this question.