Anke J. De Jong, Renze Heidstra, Herman P. Spaink, Marijke V. Hartog, Ellen A. Meijer, Theo Hendriks, Florella Lo Schiavo, Mario Terzi, Ton Bisseling, Ab Van Kammen, Sacco C. De Vries, Rhizobium Lipooligosaccharides Rescue a Carrot Somatic Embryo Mutant, The Plant Cell, Vol. 5, No. 6 (Jun., 1993), pp. 615-620
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Jong et al. (1993) studied this question.