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L. R. Cleveland, Studies on Chromosomes and Nuclear Division: Fusion of Nucleoli Independent of Chromosomal Homology; Spontaneous Aberrations, Homologous and Non-Homologous Union of Fragments; Pairing, Segregation, and Crossing-over; Photomicrographs of Living Cells during Meiotic Divisions, Transactions of the American Philosophical Society, Vol. 43, No. 3 (1953), pp. 809-869
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L. R. Cleveland (1953) studied this question.
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