Stanley R. J. Woodell, Harold A. Mooney, Harlan Lewis, The Adaptation to Serpentine Soils in California of the Annual Species Linanthus androsaceus (Polemoniaceae), Bulletin of the Torrey Botanical Club, Vol. 102, No. 5 (Sep. - Oct., 1975), pp. 232-238
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Woodell et al. (1975) studied this question.
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