Catherine Boivin, Sylvie Camut, Carlos A. Malpica, Georges Truchet, Charles Rosenberg, Rhizobium meliloti Genes Encoding Catabolism of Trigonelline Are Induced under Symbiotic Conditions, The Plant Cell, Vol. 2, No. 12 (Dec., 1990), pp. 1157-1170
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Camut et al. (1990) studied this question.