The Development of Blastocladia strangulata, N. Sp.

The plant resembles in general the other species of the genus. Its mycelium is definitely constricted, which fact, it seems, definitely places the genus in the family Leptomitaceae. 2. It possesses peculiar perforated pseudo-septa which are formed at the constrictions, and which in a way are comparable to the "cellulin rings" of other members of the Leptomitaceae. 3. Zoosporangia are provided with a number of papillae of dehiscence distributed over the surface, which are formed as the result of the gelatinization of small circular areas of the wall. The resulting plug is made up of two distinct parts, the inner of which forms a vesicle into which the zoospores escape at the time of their discharge. 4. The zoospores possess a large centrally located subtriangular mass of apparently some reserve food substance, probably proteid in nature, at whose base is located the nucleus. They are typically uniciliated, with the cilium in direct relation to the nucleus. 5. Resting sporangia possess a three-layered wall; the outer and inner layers thin and hyaline; and the middle thick, perforated, and orange colored. After a period of rest of several weeks, germination takes place by the formation of zoospores. 6. On germination the zoospore produces a germ tube which forms the basis of the rhizoid system, while the body of the spore becomes the basal cell of the plant. 7. Nuclear division is somewhat unusual, apparently, and reminds one of amitosis. It seems to the writer, however, that it is more probably a form of mitotic division dealing with a single large chromosome.