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Efrain De Luna, Angela E. Newton, Alison Withey, Dolores Gonzalez, Brent D. Mishler, The Transition to Pleurocarpy: A Phylogenetic Analysis of the Main Diplolepidous Lineages Based on rbcL Sequences and Morphology, The Bryologist, Vol. 102, No. 4 (Winter, 1999), pp. 634-650
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Luna et al. (1999) studied this question.
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