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Matthieu H. A. J. Joosten, Ralph Vogelsang, Ton J. Cozijnsen, Marianne C. Verberne, Pierre J. G. M. De Wit, The Biotrophic Fungus Cladosporium fulvum Circumvents Cf-4-Mediated Resistance by Producing Unstable AVR4 Elicitors, The Plant Cell, Vol. 9, No. 3 (Mar., 1997), pp. 367-379
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Joosten et al. (1997) studied this question.
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