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James L. Patton, Suh Y. Yang, Philip Myers, Genetic and Morphologic Divergence among Introduced Rat Populations (Rattus rattus) of the Galapagos Archipelago, Ecuador, Systematic Zoology, Vol. 24, No. 3 (Sep., 1975), pp. 296-310
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Patton et al. (1975) studied this question.
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