The inheritance of adult plant resistance to leaf rust (Puccinia recondita f. sp. tritici) was studied in the bread wheat (Triticum aestivum) cultivar Era in Morocco and the United States. Adult plant resistance was shown to be conferred by two complementary genes, Lr13 and Lr34. The magnitude of the effect resulting from the gene interaction was influenced by environment. Lr10 and an unknown gene for seedling resistance were also present in Era
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B. Ezzahiri (1989) studied this question.