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David H. Benzing, An Investigation of Two Bromeliad Myrmecophytes: Tillandsia butzii Mez, T. caput-medusae E. Morren, and Their Ants, Bulletin of the Torrey Botanical Club, Vol. 97, No. 2 (Mar. - Apr., 1970), pp. 109-115
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David H. Benzing (1970) studied this question.
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