Answers are grounded in this paper and cite their sources.
This paper develops the biometrical aspects of traits having a direct and a maternal effect. Also the influence of a grand maternal effect is considered in conjunction with the direct and maternal effect. The theoretical expectations of several common covariances between relatives are derived from a general linear formulation that involves the relationships of the contributing individuals and the relevant genic variances and covariances.
No takes yet. Share an insight, caveat, or question.
R. L. Willham (1972) studied this question.
Synapse has enriched 10 related papers. Consider them for comparative context: