Ju-Jiun Chen, David L. Mitchell, Anne B. Britt, A Light-Dependent Pathway for the Elimination of UV-Induced Pyrimidine (6-4) Pyrimidinone Photoproducts in Arabidopsis, The Plant Cell, Vol. 6, No. 9 (Sep., 1994), pp. 1311-1317
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Chen et al. (1994) studied this question.