I. L. Boyd, A. J. Woakes, P. J. Butler, R. W. Davis, T. M. Williams, Validation of Heart Rate and Doubly Labelled Water as Measures of Metabolic Rate During Swimming in California Sea Lions, Functional Ecology, Vol. 9, No. 2 (Apr., 1995), pp. 151-160
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Boyd et al. (1995) studied this question.
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