Thomas Elmqvist, Paul Alan Cox, William E. Rainey, Elizabeth D. Pierson, Restricted Pollination on Oceanic Islands: Pollination of Ceiba pentandra by Flying Foxes in Samoa, Biotropica, Vol. 24, No. 1 (Mar., 1992), pp. 15-23
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Elmqvist et al. (1992) studied this question.
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