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Sanae Kasahara, Célio F. B. Haddad, Karyotypes of Two Brazilian Microhylid Frogs of the Genus Chiasmocleis, including a New Case of Polyploidy, Journal of Herpetology, Vol. 31, No. 1 (Mar., 1997), pp. 139-142
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Kasahara et al. (1997) studied this question.
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