Carlos Bernstein, Alejandro Kacelnik, John R. Krebs, Individual Decisions and the Distribution of Predators in a Patchy Environment. II. The Influence of Travel Costs and Structure of the Environment, Journal of Animal Ecology, Vol. 60, No. 1 (Feb., 1991), pp. 205-225
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Bernstein et al. (1991) studied this question.
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