Torben Dabelsteen, Peter K. McGregor, Michael Shepherd, Xanthe Whittaker, Simon Boel Pedersen, Is the Signal Value of Overlapping Different from That of Alternating during Matched Singing in Great Tits?, Journal of Avian Biology, Vol. 27, No. 3 (Sep., 1996), pp. 189-194
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Dabelsteen et al. (1996) studied this question.