SYNOPSIS. Fitness relations in behavioral energetics can be studied using the optimality approach (cost-benefit analysis), correlational analysis of selection, the experimental approach and the comparative method, as well as other approaches. These approaches ask different questions, have different virtues and different deficiencies. By using the approaches in combination we could gain new understanding of the relationships between behavior, energy and fitness.
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Stevan J. Arnold (1988) studied this question.
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