William A. S. Sarjeant, Thurston Lacalli, Gregory Gaines, The Cysts and Skeletal Elements of Dinoflagellates: Speculations on the Ecological Causes for Their Morphology and Development, Micropaleontology, Vol. 33, No. 1 (1987), pp. 1-36
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Sarjeant et al. (1987) studied this question.
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