Willy Tinner, Brigitta Ammann, Peter Germann, Treeline Fluctuations Recorded for 12,500 Years by Soil Profiles, Pollen, and Plant Macrofossils in the Central Swiss Alps, Arctic and Alpine Research, Vol. 28, No. 2 (May, 1996), pp. 131-147
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Tinner et al. (1996) studied this question.
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