Salvador A. Morales-Verdeja, Richard C. Vogt, Terrestrial Movements in Relation to Aestivation and the Annual Reproductive Cycle of Kinosternon leucostomum, Copeia, Vol. 1997, No. 1 (Feb. 18, 1997), pp. 123-130
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Morales-Verdeja et al. (1997) studied this question.
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