^agus Nothofagus ^agus genus already differentiated in the southern Hemisphere.Northern counterparts included several species in living genera of the Betulaceae, Myricaceae, and Ulmaceae.Rapid evolution and specialion oi Nothofagus took place in the Eocene, After the pollen types and the main leaf forms were established in the Cretaceous and Eocene, respectively, the rate of morphological change was slow.More speciation was possible after the Miocene in New Guinea and also in other areas during the Plio-Pleisloccne.Several lines of evidence support former suggestions of giving Nothofagus a higher taxonomic rank.
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Edgardo J. Romero (1986) studied this question.
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