J. S. Carrion, P. Sanchez-Gomez, Palynological Data in Support of the Survival of Walnut (Juglans regia L.) in the Western Mediterranean Area During Last Glacial Times, Journal of Biogeography, Vol. 19, No. 6 (Nov., 1992), pp. 623-630
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Carrión et al. (1992) studied this question.