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Patrick J. Mulholland, Jennifer L. Tank, Diane M. Sanzone, Wilfred M. Wollheim, Bruce J. Peterson, Jackson R. Webster, Judy L. Meyer, Nitrogen Cycling in a Forest Stream Determined by a 15N Tracer Addition, Ecological Monographs, Vol. 70, No. 3 (Aug., 2000), pp. 471-493
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Mulholland et al. (2000) studied this question.
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