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Three macroalgae were studied: Enteromorpha sp. (Chlorophyceae), Hormosira banksii (Turner) Descaisne (Phaeophyceae), Corallina sp. (Rhodophyceae). Fractal dimensions (D) for the outlines of the three species were 1.24, 1.07 and 1.35 respectively in the step range 0.5–5.0 mm. All three species showed loss of epifauna on the falling tide; loss was greatest in Enteromorpha, least in Corallina. In Enteromorpha and Hormosira, most loss of epifauna was in the form of harpacticoid copepods; in Corallina loss was spread more evenly across the taxa. Epifaunal biomass and retention was much greater in Corallina than in the other two species, partially because of better water-retention properties. Enteromorpha and Hormosira both showed marked decreases in epifaunal diversity during emersion; Corallina did not.
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Davenport et al. (1999) studied this question.
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