An unusual suite of sexually dimorphic features characterizing the voiceless Bornean frog, Rana blythi, could be the product of sexual selection or intraspecific niche dimorphism. 2. Males have enlarged heads, hypertrophied jaw muscles and bony processes or fangs in the lower jaw. Males are also larger than females. 3. Alternative explanations for the sexual dimorphism are explored by testing the predictions of a biomechanical model for trophic specialization and mapping the historical transformation of the sexually dimorphic characters on a phylogeny of voiceless frogs and their relatives
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Emerson et al. (1992) studied this question.
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