An eusuchian crocodile from the Upper Cretaceous of Spain (Vilamitjana, Province of Lerida)

The first crocodilian remains from the Spanish Upper Cretaceous (Campanian–Maastrichtian) have been found in the Tremp basin (province of Lérida). Procoelous vertebrae, very strongly keeled scutes, and short stout teeth are attributed to a single adult species of an eusuchian crocodile. The Tremp crocodilian differs from all compared genera in having lumbar vertebrae with a wider neural arch and osteoderms with a tall, spine-like keel. These features, along with the general tooth morphology, suggest that the Spanish crocodile is related to the Alligatoridae.