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Michael Lynch, Michael Pfrender, Ken Spitze, Niles Lehman, Justin Hicks, Deborah Allen, Leigh Latta, Marcos Ottene, Farris Bogue, John Colbourne, The Quantitative and Molecular Genetic Architecture of a Subdivided Species, Evolution, Vol. 53, No. 1 (Feb., 1999), pp. 100-110
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Lynch et al. (1999) studied this question.
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