Answers are grounded in this paper and cite their sources.
S. P. Gittins, A. G. Parker, F. M. Slater, Population Characteristics of the Common Toad (Bufo bufo) Visiting a Breeding Site in Mid-Wales, Journal of Animal Ecology, Vol. 49, No. 1 (Feb., 1980), pp. 161-173
No takes yet. Share an insight, caveat, or question.
Gittins et al. (1980) studied this question.
Synapse has enriched 10 related papers. Consider them for comparative context: