Four-year-old ‘Marsh’ grapefruit ( Citrus paradisi Macf.) trees on trifoliate orange [ Poncirus trifoliata (L.) Raf.] rootstock were temperature acclimated to 5°C in controlled environment facilities with approximately 400 μeinsteins m −2 s −1 PAR. Total soluble carbohydrates and proline increased in both leaves and fruit flavedo as temperatures were progressively decreased. Maximum accumulation of carbohydrates occurred in leaves and flavedo at 10° ambient air. Both sucrose and reducing sugars increased in leaves at all acclimating temperatures, but only reducing sugars increased in the flavedo at temperatures below 15°. The concentration of proline was the greatest in the leaves and flavedo at 5°. Both total soluble carbohydrates and proline decreased during temperature deacclimation at 25°.
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Purvis et al. (1982) studied this question.