Peter A. Anderson, Gregory J. Lawrence, Bronwyn C. Morrish, Michael A. Ayliffe, E. Jean Finnegan, Jeffrey G. Ellis, Inactivation of the Flax Rust Resistance Gene M Associated with Loss of a Repeated Unit within the Leucine-Rich Repeat Coding Region, The Plant Cell, Vol. 9, No. 4 (Apr., 1997), pp. 641-651
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Anderson et al. (1997) studied this question.